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Browse subject: mixed forests
Number of records: 120

Detection Probabilities of Woodpecker Nests in Mixed Conifer Forests in Oregon
About this Resource: Accurate estimates of Black-backed (Picoides arcticus) and Hairy Woodpecker (P. villosus) nests and nest survival rates in post-fire landscapes provide land managers with information on the relative importance of burned forests to nesting woodpeckers. We conducted multiple-observer surveys in burned and unburned mixed coniferous forests in Oregon to identify important factors influencing detection rates of woodpecker nests. We found 21 Black-backed Woodpecker nests and 38 Hairy Woodpecker nests in burned forest, and three Hairy Woodpecker nests in unburned forest. Competing models of detection probability in Program MARK indicated that nest-detection probability differed by nest stage. We found no evidence to indicate that detection rates of nests were associated with survey timing during the nesting season. Raw nest counts in burned coniferous forests may underestimate nest numbers, especially for nests in early stages of development. Black-backed Woodpecker nests were slightly more detectable than those of Hairy Woodpeckers in burned forests, and observers may differ in their abilities to detect nests.

Microhabitat Associations of Northern Flying Squirrels in Burned and Thinned Forest Stands of the Sierra Nevada
About this Resource: Prescribed burning and mechanical thinning are used to manage fuels within many western North American forest ecosystems, but few studies have examined the relative impacts of these treatments on forest wildlife. We sampled northern flying squirrels (Glaucomys sabrinus) and microhabitat variables in burned, thinned and control stands of mixed-conifer forest of the southern Sierra Nevada at the Teakettle Experimental Forest. We used this information to determine the effects of burning and thinning on the microhabitat associations of flying squirrels. Across pretreatment stands, the probability of flying squirrel capture increased with decreasing distance to a perennial creek and increasing litter depth. The probability of flying squirrel capture also was greater with increased canopy cover in thinned stands and increased litter depth in burned stands. Greater canopy cover may provide protection from predators, thicker litter layers may harbor a greater abundance of truffles, a primary food of northern flying squirrels, and creeks may provide squirrels with food sources, drinking water and nest trees. Results from this study underscore the need for more information on the effects of forest management on northern flying squirrels near the southern extent of the species' geographic range.

Prescribed fire effects on the herbaceous layer of mixed-oak forests.
About this Resource: In 1994, a multidisciplinary project was established to study the effects of prescribed fire on oak forests in southern Ohio. Here we describe the herbaceous layer response to fires over a 5-year period. In four study sites, treatments imposed were unburned, periodic (1996 and 1999), and annual (1996-1999) fires. Sample plots (n = 108) were stratified by an integrated moisture index. Species' frequencies were recorded annually, and a total of 452 species (97% native) were documented. Though species composition was significantly affected by fire, the effects were shown by ordination to be small in magnitude relative to overall compositional variation. Burned areas developed greater small-scale species richness as grasses, summer forbs, and seed-banking species increased in frequency; however, these changes were also not large in magnitude. Though a few species increased substantially via germination after fire, most common species exhibited frequency increases or decreases of <10% on burned units. Fire effects on vegetation were largely similar between annual and periodic burns and also among integrated moisture index classes. Direct fire effects on vegetation were limited by the dormant-season timing of burns and the resprouting of woody plants. Indirect effects were limited, as fires caused relatively minor changes in forest structure and resource availability in these long-unburned forests.

Comparisons of land cover and LAI estimates derived from ETM+ and MODIS for four sites in North America: a quality assessment of 2000/2001 provisional MODIS products
About this Resource: The MODIS land science team produces a number of standard products, including land cover and leaf area index (LAI). Critical to the success of MODIS and other sensor products is an independent evaluation of product quality. In that context, we describe a study using field data and Landsat ETM+ to map land cover and LAI at four 49-km2 sites in North America containing agricultural cropland (AGRO), prairie grassland (KONZ), boreal needleleaf forest, and temperate mixed forest. The purpose was to: (1) develop accurate maps of land cover, based on the MODIS IGBP (International Geosphere-Biosphere Programme) land cover classification scheme; (2) derive continuous surfaces of LAI that capture the mean and variability of the LAI field measurements; and (3) conduct initial MODIS validation exercises to assess the quality of early (i.e., provisional) MODIS products. ETM+ land cover maps varied in overall accuracy from 81% to 95%. The boreal forest was the most spatially complex, had the greatest number of classes, and the lowest accuracy. The intensive agricultural cropland had the simplest spatial structure, the least number of classes, and the highest overall accuracy. At each site, mapped LAI patterns generally followed patterns of land cover across the site. Predicted versus observed LAI indicated a high degree of correspondence between field-based measures and ETM+ predictions of LAI. Direct comparisons of ETM+ land cover maps with Collection 3 MODIS cover maps revealed several important distinctions and similarities. One obvious difference was associated with image/map resolution. ETM+ captured much of the spatial complexity of land cover at the sites. In contrast, the relatively coarse resolution of MODIS did not allow for that level of spatial detail. Over the extent of all sites, the greatest difference was an overprediction by MODIS of evergreen needleleaf forest cover at the boreal forest site, which consisted largely of open shrubland, woody savanna, and savanna. At the agricultural, temperate mixed forest, and prairie grassland sites, ETM+ and MODIS cover estimates were similar. Collection 3 MODIS-based LAI estimates were considerably higher (up to 4 m2 m-2) than those based on ETM+ LAI at each site. There are numerous probable reasons for this, the most important being the algorithms' sensitivity to MODIS reflectance calibration, its use of a prelaunch AVHRR-based land cover map, and its apparent reliance on mainly red and near-IR reflectance. Samples of Collection 4 LAI products were examined and found to consist of significantly improved LAI predictions for KONZ, and to some extent for AGRO, but not for the other two sites. In this study, we demonstrate that MODIS reflectance data are highly correlated with LAI across three study sites, with relationships increasing in strength from 500 to 1000 m spatial resolution, when shortwave-infrared bands are included.

Soil respiration response to prescribed burning and thinning in mixed-conifer and hardwood forests.
About this Resource: The effects of management on soil carbon efflux in different ecosystems are still largely unknown yet crucial to both our understanding and management of global carbon flux. To compare the effects of common forest management practices on soil carbon cycling, we measured soil respiration rate (SRR) in a mixed-conifer and hardwood forest that had undergone various treatments from June to August 2003. The mixed-conifer forest, located in the Sierra Nevada Mountains of California, had been treated with thinning and burning manipulations in 2001, and the hardwood forest, located in the southeastern Missouri Ozarks, had been treated with harvesting manipulations in 1996 and 1997. Litter depth, soil temperature, and soil moisture were also measured. We found that selective thinning produced a similar effect on both forests by elevating SRR, soil moisture, and soil temperature, although the magnitude of response was greater in the mixed-conifer forest. Selective harvest increased SRR by 43% (from 3.38 to 4.82 micromol.m(-2).s(-1)) in the mixed-conifer forest and by 14% (from 4.25 to 4.84 micromol.m(-2).s(-1)) in the hardwood forest. Burning at the conifer site and even-aged harvesting at the mixed-hardwood site did not produce significantly different SRR from controls. Mean SRR were 3.24, 3.42, and 4.52 micromol.m(-2).s(-1), respectively. At both sites, manipulations did significantly alter SRR by changing litter depth, soil structure, and forest microclimate. SRR response varied by vegetation patch type, the scale at which treatments altered these biotic factors. Our findings provide forest managers first-hand information on the response of soil carbon efflux to various management strategies in different forests.

An Evaluation of the Contribution of Cultivated Allspice (Pimenta Dioca) to Vertebrate Biodiversity Conservation in Nicaragua
About this Resource: Tropical deforestation has emerged as one of the most important conservation challenges of our time, both because of the high species diversity and rates of endemism of tropical forests, and because of the rapid rate at which this process is proceeding. Recent studies indicate that areas of low-intensity agroforestry have similar levels of vertebrate diversity as some primary habitats, leading some researchers and conservationists to conclude that this type of commodity production could contribute to the conservation of biodiversity. We compared the composition of bird, mammal and herpetofaunal communities in primary forest, secondary forest, and pasture--and within the allspice productive systems that have replaced pasture. We found that mammal species richness was higher in primary forest than all other habitats; however for resident and migrant birds, amphibians and reptiles, species richness was similar between primary forest and the other habitats. Despite similarities in overall numbers of species, there were numerous species that were encountered only in primary habitats. We conclude that the cultivation of allspice in a mixed productive system can offset some of the losses to biodiversity; however it should be complemented by the establishment and maintenance of protected areas to accommodate populations of primary forest specialists that are unable to persist in altered habitats.

Testing IsoSource: stable isotope analysis of a tropical fishery with diverse organic matter sources.
About this Resource: We sampled consumers and organic matter sources (mangrove litter, freshwater swamp-forest litter, seagrasses, seagrass epiphytes, and marine particulate organic matter [MPOM]) from four estuaries on Kosrae, Federated States of Micronesia for stable isotope (sigma13C and sigma34S) analysis. Unique mixing solutions cannot be calculated in a dual-isotope, five-endmember scenario, so we tested IsoSource, a recently developed statistical procedure that calculates ranges in source contributions (i.e., minimum and maximum possible). Relatively high minimum contributions indicate significant sources, while low maxima indicate otherwise. Litter from the two forest types was isotopically distinguishable but had low average minimum contributions (0-8% for mangrove litter and 0% for swamp-forest litter among estuaries). Minimum contribution of MPOM was also low, averaging 0-13% among estuaries. Instead, local marine sources dominated contributions to consumers. Minimum contributions of seagrasses averaged 8-47% among estuaries (range 0-88% among species). Minimum contributions of seagrass epiphytes averaged 5-27% among estuaries (range 0-69% among species). IsoSource enabled inclusion of five organic matter sources in our dual-isotope analysis, ranking trophic importance as follows: seagrasses > seagrass epiphytes > MPOM > mangrove forest > freshwater swamp-forest. IsoSource is thus a useful step toward understanding which of multiple organic matter sources support food webs; more detailed work is necessary to identify unique solutions.

Predation of gypsy moth (Lepidoptera: Lymantriidae) pupae in three ecosystems along the southern edge of infestation
About this Resource: The predation potential of small mammals, in particular mice, Peromyscus spp., and invertebrates, was evaluated from 1992 to 1995 near the leading edge of gypsy moth, Lymantria dispar (L.), spread into the southeastern United States. Two study sites were established in each of three geographic areas: the coastal plain, Piedmont, and mountains. All sites were mixed hardwood stands with varying amounts of oak, Quercus spp., and all were classified for gypsy moth susceptibility. Small mammal density was estimated using Sherman live-traps and pitfall traps within these 4.68-ha sites in early and late summer. Each site contained 75 trapping stations located on a 25-m grid. Predation was measured by offering freeze-dried gypsy moth pupae near trapping stations at four heights (0, 0.25, 1.0, and 2.0 m) on different tree boles. Pupal predation was monitored for three consecutive nights. Vertebrate predation was positively correlated with good mast production in the previous autumn. Predation data showed that when mice were at high densities they were the major source of pupal predation. However, within these southern sites, when densities of Peromyscus spp. were low, predation by invertebrates was occasionally greater than predation by vertebrates. These data suggest that in some years invertebrates may retard gypsy moth buildup when small mammals are scarce due to mast crop failures.

Carbon sequestration by forests and soils on mined land in the Midwestern and Appalachian coalfields of the U.S
About this Resource: Carbon (C) accreditation of forest development projects is one approach for sequestering atmospheric CO₂, under the provisions of the Kyoto protocol. The C sequestration potential of reforested mined land is not well known. The purpose of this work was to estimate and compare the ecosystem C content in forests established on surface, coal-mined and non-mined land. We used existing tree, litter, and soil C data for fourteen mined and eight adjacent, non-mined forests in the Midwestern and Appalachian coalfields to determine the C sequestration potential of mined land reclaimed prior to the passage of the Surface Mining Control and Reclamation Act (1977). We developed statistically significant and biologically reasonable models for ecosystem C across the spectrum of site quality and stand age. On average, the highest amount of ecosystem C on mined land was sequestered in pine stands (148Mgha⁻¹), followed by hardwood (130Mgha⁻¹) and mixed stands (118Mgha⁻¹). Non-mined hardwood stands sequestered 210MgCha⁻¹, which was about 62% higher than the average of all mined stands. Our mined land response surface models of C sequestration as a function of site quality and age explained 59, 39, and 36% of the variation of ecosystem C in mixed, pine, and hardwood stands, respectively. In pine and mixed stands, ecosystem C increased exponentially with the increase of site quality, but decreased with age. In mined hardwood stands, ecosystem C increased asymptotically with age, but it was not affected by site quality. At rotation age (60 yr), ecosystem C in mined hardwood stands was less on high quality sites, but similar for low quality sites compared to non-mined hardwood stands. The overall results indicated that the higher the original forest site quality, the less likely C sequestration potential was restored, and the greater the disparity between pre- and post-mining C sequestration stocks.

Survival of larvae and nymphs of Ixodes scapularis say (Acari: Ixodidae) in four habitats in Maryland.
About this Resource: Host-seeking and fed larvae and nymphs of the blacklegged tick, Ixodes scapularis Say, were placed in mesh packets and in vials in the leaf litter on the floor of mixed deciduous forest, Virginia pine-southern red oak forest, and white pine plantations with and without Nepal microstegium, Microstegium vimineum (Trinius) A. Camus. An introduced shade-tolerant grass, Nepal microstegium, is expanding its range northeastward into areas densely populated with I. scapularis. As determined by flag sampling, the density of host-seeking nymphs at the Virginia pine sites was much lower than in the other habitats. None of the four habitats appeared to be consistently more favorable or unfavorable for the survival of confined fed and unfed I. scapularis larvae and nymphs. More unfed nymphs survived in vials than in packets in Virginia pine and white pine with Nepal grass sites. Fed larvae and nymphs tended to survive the summer better than unfed ticks.

A survey for nymphs of host-seeking Ixodes scapularis and Amblyomma americanum (Acari: Ixodidae) on tree trunks in deciduous forests.
About this Resource: Trunks of 83 trees in a mixed deciduous forests in Maryland were sampled for the presence of nymphs of the blacklegged tick, Ixodes scapularis Say, and the lone star tick, Amblyomma americanum (L.). Although one or more nymphs of either I. scapularis or A. americanum was found in leaf litter and substrate less than or equal to 1 m from the bases of 47% of the trees sampled, a total of 6 I. scapularis nymphs was found on the trunks of only five trees. No nymphs were found on the trunks of 12 dead trees. No A. americanum nymphs were found on any tree trunks. The trunks were sampled to 2.5 m above the soil, but the nymphs were found less than or equal to 1 m from the ground. More than 50% of I. scapularis nymphs found in the leaf litter less than or equal to 1 m from bases of living trees were north of the trees sampled, whereas few I. scapularis were found west of trees. These findings suggest that the I. scapularis nymphs' presence on tree trunks is of little ecological consequence, unless nymphs were being removed from tree trunks by acquiring hosts at such a rapid rate that nymphal numbers on trunks could not accrue.

Patterns of growth dominance in forests of the Rocky Mountains, USA
About this Resource: We used data from 142 stands in Colorado and Wyoming, USA, to test the expectations of a model of growth dominance and stand development. Growth dominance relates the distribution of growth rates of individual trees within a stand to tree sizes. Stands with large trees that account for a greater share of stand growth than of stand mass exhibit strong growth dominance. Stands with large trees that contribute less to stand growth than to stand mass show reverse growth dominance. The four-phase model predicts that forests move from a period of little dominance (Phase 1), with trees accounting for similar contributions to stand growth and stand mass. Phase 2 is a period of strong growth dominance, where larger trees account for a disproportionately large amount of total stand growth. Growth dominance declines during Phase 3 as growth of the larger trees slows. A final Phase 4 shows reverse growth dominance when the growth of larger trees is less than their proportional contribution to total stand mass. The datasets supported the expectation of reverse growth dominance in old forests of ponderosa pine, Engelmann spruce and subalpine fir, lodgepole pine, and mixed stands of aspen and conifers. Pure aspen stands did not show reverse growth dominance. An age sequence of lodgepole pine failed to show the expected Phase 2 period of strongly developed growth dominance. Future work needs to combine quantitative descriptions of patterns in growth dominance with experimental manipulations of resource supplies and environmental conditions to connect forest dynamics at the scales of individual trees, groups of trees, and stands.

Influence of Cow Age on Grazing Distribution in a Mixed-Conifer Forest
About this Resource: Optimal distribution of cattle on forested rangelands has long been a subject of concern specifically related to uniform and sustainable use of forage resources. Our objective was to determine if cow age influenced distribution and resource use on forested rangelands. This study was conducted from 1991 to 2001 at the US Department of Agriculture Starkey Experimental Forest and Range, northeastern Oregon, a mixed-conifer forested rangeland. We used 43039 locations of cattle taken from 1 h prior to sunrise until 4 h after sunrise and 4 h prior to sundown until 1 h after sundown from 15 July to 30 August to evaluate cattle distribution patterns during peak foraging time. Cattle were grouped into four age classes: 2- and 3-yr-old cattle, 4- and 5-yr-old cattle, 6- and 7-yr-old cattle, and cattle >=8 yr old. All age classes preferred areas with gentler slopes (P<0.05), westerly aspects (P<0.05), farther from water (P<0.05), and with greater forage production (P<0.05) than pasture averages. Cattle older than 3 yr of age selected areas with less canopy closure (P<0.05) than the mean value for the pasture. Young cows (<4 yr old) selected lower elevations and steeper slopes than the oldest cows (P<0.05). In summary, cow age and correspondingly its experience directly influences distribution patterns and forage resource use of cattle at the Starkey Experimental Forest and Range.

Transmission of Phytophthora ramorum in mixed-evergreen forest in California.
About this Resource: During 2001 to 2003, the transmission biology of Phytophthora ramorum, the causal agent of sudden oak death, was studied in mixed-evergreen forest, a common forest type in northern, coastal California. Investigation of the sources of spore production focused on coast live oak (Quercus agrifolia) and bay laurel (Umbellularia californica), dominant hosts that comprised 39.7 and 46.2% of the individuals at the study site, respectively. All tests for inoculum production from the surface of infected coast live oak bark or exudates from cankers were negative. In contrast, sporangia and chlamydospores were produced on the surface of infected bay laurel leaves. Mean number of zoospores produced from infected bay laurel leaves under natural field conditions during rainstorms was 1,173.0 +/- SE 301.48, and ranged as high as 5,200 spores/leaf. P. ramorum was recovered from rainwater, soil, litter, and streamwater during the mid- to late rainy season in all 3 years of the study. P. ramorum was not recovered from sporadic summer rains or soil and litter during the hot, dry summer months. Concentrations of inoculum in rainwater varied significantly from year to year and increased as the rainy season progressed for the two complete seasons that were studied. Potential dispersal distances were investigated for rainwater, soil, and streamwater. In rainwater, inoculum moved 5 and 10 m from the inoculum source. For soil, transmission of inoculum was demonstrated from infested soil to bay laurel green leaf litter, and from bay laurel green leaf litter to aerial leaves of bay laurel seedlings. One-third to one-half of the hikers tested at the study site during the rainy season also were carrying infested soil on their shoes. In streamwater, P. ramorum was recovered from an unforested site in pasture approximately equal to 1 km downstream of forest with inoculum sources. In total, these studies provide details on the production and spread of P. ramorum inoculum in mixed-evergreen forest to aid forecasting and managing disease transmission of this environmentally destructive pathogen.

Snag longevity under alternative silvicultural regimes in mixed-species forests of central Maine.
About this Resource: Predictions of snag longevity, defined here as the probability of snag survival to a given age, are key to designing silvicultural regimes that ensure their availability for wildlife and form an important component of carbon flow models. Species, diameter at breast height, stand density, management regime, and agent of tree mortality were assessed for their effect on snag longevity in a long-term silvicultural study on the Penobscot Experimental Forest in central Maine. Snag recruitment and fall data from USDA Forest Service inventories between 1981 and 1997 were analyzed using parametric survival analysis. A Weibull model fit the data best, indicating a significant lag time followed by rapid fall rates. Half-times varied among species, with Thuja occidentalis L. having the longest (10 years) and Picea species the shortest (6 years). Snag longevity was significantly greater with increasing diameter and decreased with increasing stand density. Agent of mortality and silvicultural treatment were also significant. Two models were developed for estimating probability of snag survival over time, one that included predictor variables unique to the silvicultural systems study on the Penobscot Experimental Forest and one using predictor variables available in most standard inventories. Snag survival models can easily be incorporated into comprehensive forest dynamics models to facilitate estimates of wildlife habitat structure and carbon flow.

Nitrogen mineralization and nitrification in a mixed-conifer forest in southern California: controlling factors, fluxes, and nitrogen fertilization response at a high and low nitrogen deposition site.
About this Resource: Net fluxes of nitrogen (N) mineralization and nitrification were measured in situ on a monthly basis for 3 years at a high (HN) and low (LN) N deposition site in the San Bernardino Mountains, California. Mean N mineralization fluxes in the forest floor and top 10 cm of mineral soil were 19.0 and 59.8 kg N.ha(-1).year(-1) at LN and HN, respectively. Mean net nitrification fluxes were 11.2 and 55.9 kg N.ha(-1).year(-1) at LN and HN, respectively. Relative nitrification (the percent N mineralized that was nitrified) was generally lower under Pinus ponderosa Dougl. ex P. & C. Laws. (or Pinus jeffreyi Grev. & Balf.) canopies than under Quercus kelloggii Newb. or open canopies. The rate of net N mineralization was the key factor for predicting the rate of net nitrification. Fertilization with 50 and 150 kg N.ha(-1) at LN significantly increased the rates of net mineralization and net nitrification. At HN fertilization had no significant effect on net nitrification. We conclude that at low-deposition sites increased nitrification occurs in the short term in response to added N, but that sustained elevated net nitrification is driven by the accumulation of N-enriched litter and soil organic matter in conjunction with chronic throughfall N deposition inputs.

A Comparison of Statistical Methods for Estimating Forest Biomass from Light Detection and Ranging Data.
About this Resource: Strong regression relationships between light detection and ranging (LIDAR) metrics and indices of forest structure have been reported in the literature. However, most papers focus on empirical results and do not consider LIDAR metric selection and biological interpretation explicitly. In this study, three different variable selection methods (stepwise regression, principle component analysis [PCA], and Bayesian modeling averaging [BMA]) were compared using LIDAR data from three study sites: Capitol Forest in western Washington State, Mission Creek in central Washington State, and Kenai Peninsula in south central Alaska. Separate aboveground biomass regression models were developed for each site as well as common models using three study sites simultaneously. Final biomass models have R 2 values ranging from 0.67 to 0.88 for three study sites. PCA indicates that three LIDAR metrics (mean height, coefficient variation of height, and canopy LIDAR point density) explain the majority of variation contained within a larger set of metrics. Within each study area, forest biomass models using these three predictor variables had similar R 2 values as the stepwise and BMA regression models. Individual site models using these three variables are recommended because these models are straightforward in terms of model form and biological interpretation and are easily adopted for application.

Development of equations for predicting Puerto Rican subtropical dry forest biomass and volume
About this Resource: Carbon accounting, forest health monitoring and sustainable management of the subtropical dry forests of Puerto Rico and other Caribbean Islands require an accurate assessment of forest aboveground biomass (AGB) and stem volume. One means of improving assessment accuracy is the development of predictive equations derived from locally collected data. Forest inventory and analysis (FIA) measured tree diameter and height, and then destructively sampled 30 trees from 6 species at an upland deciduous dry forest site near Ponce, Puerto Rico. This data was used to develop best parsimonious equations fit with ordinary least squares procedures and additive models fit with nonlinear seemingly unrelated regressions that estimate subtropical dry forest leaf, woody, and total AGB for Bucida buceras and mixed dry forest species. We also fit equations for estimating inside and outside bark total and merchantable stem volume using both diameter at breast height (d.b.h.) and total height, and diameter at breast height alone for B. buceras and Bursera simaruba. Model fits for total and woody biomass were generally good, while leaf biomass showed more variation, possibly due to seasonal leaf loss at the time of sampling. While the distribution of total AGB into components appeared to remain relatively constant across diameter classes, AGB variability increased and B. simaruba and B. buceras allocated more carbon into branch biomass than the other species. When comparing our observed and predicted values to other published dry forest AGB equations, the equation developed in Mexico and recommended for areas with rainfall >900 mm/year gave estimates substantially lower than our observed values, while equations developed using dry forest data from forest in Australia, India and Mexico were lower than our observed values for trees with d.b.h. <25 cm and slightly higher for trees with d.b.h. >30 cm. Although our ability to accurately estimate merchantable stem volume and live tree AGB for subtropical dry forests in Puerto Rico and other Caribbean islands has been improved, much work remains to be done to sample a wider range of species and tree sizes.

Long-term development of above- and below-ground carbon stocks following land-use change in subalpine ecosystems of the Swiss National Park
About this Resource: Vegetation changes following agricultural land abandonment at high elevation --- which is frequent in Europe --- could have a major impact on carbon (C) sequestration. However, most information on the effects of vegetation changes on ecosystem C stocks originates from low-elevation studies on reforestation or early successional forests, and little is known about how these stocks change during long-term secondary forest succession and at high elevation. We assessed aboveground, belowground, and ecosystem organic matter and C stocks in high-elevation ecosystems that represent the long-term development (centuries) following land abandonment: short- and tall-grass pastures, Swiss mountain pine (Pinus mugo Turra), mixed-conifer, and Swiss stone pine (Pinus cembra L.) -- European larch (Larix decidua P. Mill.) forests. Aboveground C stocks were lowest in the short-grass pastures (0.1 Mg C·ha-1) and reached a maximum in the mixed-conifer and stone pine - larch forests (166 Mg C·ha-1). Belowground C stocks did not differ among the ecosystems studied. We only detected ecosystem C sequestration during reforestation; whereas no significant differences in ecosystem C stocks were found during long-term secondary forest development. Our calculations showed that only an additional 1733-3032 Mg C·year-1 would be sequestered owing to natural reforestation in high-elevation Switzerland, which likely can be considered negligible compared with total annual C sequestration calculated for Swiss forests in other studies.

Short-term effects of fire and forest thinning on truffle abundance and consumption by Neotamias speciosus in the Sierra Nevada of California.
About this Resource: In many western North American forests, prescribed burning and mechanical thinning are widely used to reduce fuels and restore stand conditions after a century of fire suppression. Few studies have followed the relative impacts of these treatments on the production and consumption of truffles in forest ecosystems, particularly in the Sierra Nevada of California. Using a full-factorial completely randomized design, we examined the short-term impacts of prescribed burning (no burn and burn), mechanical thinning (no thin, light thin, and heavy thin), and combinations of these treatments on the production of truffles and their consumption by lodgepole chipmunks (Neotamias speciosus Merriam) in a mixed-conifer forest of the southern Sierra Nevada of California. Truffle frequency, biomass, and species richness were lower in thinned or burned plots than controls, as was the frequency and generic richness of truffles in the diet of N. speciosus. Truffle frequency, biomass, and species richness, and truffle consumption by N. speciosus were lower in heavily thinned and thinned and burned plots than in those exclusively burned. These results suggest that either thinning or burning can reduce short-term truffle production and consumption, and potentially the dispersal of ectomycorrhizal spores by small mammals. Moreover, truffles decreased with treatment intensity, suggesting heavy thinning and higher burn intensity, particularly when applied together, can significantly affect short-term truffle abundance and small mammal consumption.

Assessing the effects of vegetation types on carbon storage fifteen years after reforestation on a Chinese fir site
About this Resource: Forest ecosystems play a significant role in sequestering carbon (C) in biomass and soils. Plantations established in subtropical China since the 1980s, mainly of Chinese fir (Cunninghamia lanceolata (Lamb.) Hook) in monocultures, have proved to be major C sinks. However, information is lacking about whether mixing Chinese fir with broadleaved tree species will increase stand growth and C sequestration. We address this question by comparing a pure Chinese fir plantation and two mixed plantations established in 1990 at Huitong Experimental Station of Forest Ecology, Hunan Province, China. The mixed plantations include Chinese fir and either Kalopanax septemlobus (Thunb.) Koidz or Alnus cremastogyne Burk., planted at 4:1 ratios. We found that total C storage was 123, 131 and 142Mgha⁻¹ in the pure plantation, mixed plantation with K. septemlobus, and mixed plantation with A. cremastogyne, respectively. The mixed plantation with A. cremastogyne increased C storage in biomass relative to the pure Chinese fir plantation (P <0.05). No significant difference was detected between mixed plantations. Soil C storage did not differ among these plantations, ranging from 67.9±7.1 to 73.3±9.1Mgha⁻¹, which accounted for about 55% of the total C pools. Our results indicated that as the mixture of Chinese fir and broadleaved species will increase both biomass C and soil C storage over pure Chinese fir, and will do it, within 15 years of planting.

Effects of Natural Barriers and Habitat on the Western Spread of Raccoon Rabies in Alabama.
About this Resource: Although domestic animal transmission of rabies has largely been mitigated, the disease remains a concern in both Europe and North America where wildlife transmission has caused epizootics. Raccoon (Procyon lotor) rabies was established in Alabama, USA, in 1975, primarily in the southeastern corner of the state. However, with the exception of isolated events, rabies has not continued to spread westward across the Alabama River. We monitored movements of 100 radiocollared raccoons on 2 sites within hardwood and agriculture habitats in a rabies enzootic area east of the Alabama River, in managed pine habitat area west of the river where rabies sporadically occurs, and in a mixed pine hardwood area outside of the known rabies enzootic area to determine if raccoon movements and habitat use in certain habitat types and the presence of a river may serve as natural barriers preventing the western spread of rabies in Alabama. We also examined raccoon contact rates to determine if they influence disease transmission through static and dynamic interactions. Raccoons in mixed pine-hardwood forest habitats had smaller home ranges and less overlap of ranges compared to the other 3 habitats. However, static interactions between habitats in the use of overlap areas did not differ (F11,129 = 1.63, P = 0.09). Rabies antibody titers were highest in the managed pine habitat (28%) even prior to oral vaccine bait distributions in spring of 2004 and 2005. Biomarker data from radiocollared and additional raccoons captured after the bait distribution west of the Alabama River demonstrated a low efficacy of the vaccine reaching the small southern raccoons. The combination of the river as a partial barrier, the high percentage of pine forested habitat west of the river, and limited spatial movements of raccoons within these forested habitats appears to have reduced the likelihood of rabies establishing west of the river. Understanding different host-habitat-disease systems is important for successful management of diseases. Based on our results, we recommend that the oral vaccine program continue to use the Alabama River as a partial barrier and baiting be concentrated in the fragmented bottomland hardwood forests and around larger bodies of water where raccoon densities are highest. Success of baiting strategies designed to take advantage of northern raccoon dynamics and habitat use may not be applicable to southern populations.

Influence of prescribed fire on carabid beetle (Carabidae) and spider (Araneae) assemblages in forest litter in southwestern Oregon
About this Resource: The objective of this study was to determine if prescribed fire affects spider (Araneae) and carabid beetle (Carabidae) abundance, and whether the magnitude of this effect varies with time since fire. Within mixed conifer stands, nine understory fuels-reduction burns, ranging from <1 to 15 yr old, were compared with adjacent unburned sites. Pitfall traps were used to compare macroarthropod abundance over 5 mo. In total, 3,441 spiders in 24 families and >120 species, and 14,793 carabid beetles from 17 species, were identified from the samples. Seven spider families and five species of carabid beetles were abundant enough to be analyzed statistically. Four spider families were more abundant in unburned sites (Antrodiaetidae, Cybaeidae, Thomisidae and Linyphiidae) while three families were more numerous in burned sites (Lycosidae, Gnaphosidae and Dictynidae). Four of five carabid beetle species were more abundant in unburned sites [Pterostichus herculaneus Mannerheim, P. setosus Hatch, Scaphinotus rugiceps rugiceps (Horn) and Zacotus matthewsii LeConte]. There was no difference found for Omus cazieri van den Berghe. No differences in species richness or diversity (Simpson, Shannon-Wiener and Berger-Parker indices) were found for spiders or carabid beetles. Overall, the relationship between abundance and time since burning was weak, with marginal significance found only for Dictynidae and Gnaphosidae. We suggest that changes in foraging substrate, prey availability or microclimatic conditions since fire may have interacted with life history characteristics to influence the abundance of these organisms. Differences in fire intensities among years may have masked patterns in arthropod abundance associated with time since burning.

Photosynthetic characteristics of Fagus sylvatica and Quercus robur established for stand conversion from Picea abies
About this Resource: Efforts in Europe to convert Norway spruce (Picea abies) plantations to broadleaf or mixed broadleaf-conifer forests could be bolstered by an increased understanding of how artificial regeneration acclimates and functions under a range of Norway spruce stand conditions. We studied foliage characteristics and leaf-level photosynthesis on 7-year-old European beech (Fagus sylvatica) and pedunculate oak (Quercus robur) regeneration established in open patches and shelterwoods of a partially harvested Norway spruce plantation in southwestern Sweden. Both species exhibited morphological plasticity at the leaf level by developing leaf blades in patches with an average mass per unit area (LMA) 54% greater than of those in shelterwoods, and at the plant level by maintaining a leaf area ratio (LAR) in shelterwoods that was 78% greater than in patches. However, we observed interspecific differences in photosynthetic capacity relative to spruce canopy openness. Photosynthetic capacity (A ₁₆₀₀, net photosynthesis at a photosynthetic photon flux density of 1600μmolphotonsm⁻² s⁻¹) of beech in respect to the canopy gradient was best related to leaf mass, and declined substantially with increasing canopy openness primarily because leaf nitrogen (N) in this species decreased about 0.9mgg⁻¹ with each 10% rise in canopy openness. In contrast, A ₁₆₀₀ of oak showed a weak response to mass-based N, and furthermore the percentage of N remained constant in oak leaf tissues across the canopy gradient. Therefore, oak photosynthetic capacity along the canopy gradient was best related to leaf area, and increased as the spruce canopy thinned primarily because LMA rose 8.6gm⁻² for each 10% increase in canopy openness. These findings support the premise that spruce stand structure regulates photosynthetic capacity of beech through processes that determine N status of this species; leaf N (mass basis) was greatest under relatively closed spruce canopies where leaves apparently acclimate by enhancing light harvesting mechanisms. Spruce stand structure regulates photosynthetic capacity of oak through processes that control LMA; LMA was greatest under open spruce canopies of high light availability where leaves apparently acclimate by enhancing CO₂ fixation mechanisms.

Radiation use efficiency in adjacent hardwood and pine forests in the southern Appalachians
About this Resource: The efficiency with which trees convert photosynthetically active radiation (PAR) to biomass has been shown to be consistent within stands of an individual species, which is useful for estimating biomass production and carbon accumulation. However, radiation use efficiency (ε) has rarely been measured in mixed-species forests, and it is unclear how species diversity may affect the consistency of ε, particularly across environmental gradients. We compared aboveground net primary productivity (ANPP), intercepted photosynthetically active solar radiation (IPAR), and radiation use efficiency (ε =ANPP/IPAR) between a mixed deciduous forest and a 50-year-old white pine (Pinus strobus L.) plantation in the southern Appalachian Mountains. Average ANPP was similar in the deciduous forest (11.5Mgha⁻¹ y⁻¹) and pine plantation (10.2Mgha⁻¹ y⁻¹), while ε was significantly greater in the deciduous forest (1.25gMJ⁻¹) than in the white pine plantation (0.63gMJ⁻¹). Our results demonstrate that late-secondary hardwood forests can attain similar ANPP as mature P. strobus plantations in the southern Appalachians, despite substantially less annual IPAR and mineral-nitrogen availability, suggesting greater resource-use efficiency and potential for long-term carbon accumulation in biomass. Along a 260m elevation gradient within each forest there was not significant variation in ε. Radiation use efficiency may be stable for specific forest types across a range of environmental conditions in the southern Appalachian Mountains, and thus useful for generating estimates of ANPP at the scale of individual watersheds.

Effects of alternative silviculture on stump sprouting in the southern Appalachians
About this Resource: Stump sprouts are an important form of regeneration for a number of species in the southern Appalachians, especially the oaks (Quercus spp.). Alternative regeneration systems to clearcutting such as shelterwood and leave-tree systems are being implemented in many hardwood stands in the Appalachians. However, the effects of these alternative silvicultural systems on stump sprouts are not known. Therefore, we evaluated the impact of three silvicultural systems: a clearcut, leave-tree, and shelterwood on stump sprouting. These treatments were implemented in seven stands in Virginia and West Virginia in the Appalachian Plateau (AP) and Ridge and Valley (RV) physiographic provinces. The stands were even-aged oak dominated Appalachian hardwood stands with ages ranging from 62 to 100 years. Species were placed into six groups: (1) red oak (Quercus spp.), (2) chestnut oak (Q. prinus L.), (3) white oak (Q. alba L.) and hickory (Carya spp.), (4) red maple (Acer rubrum L.), (5) mixed mesic, and (6) midstory groups. Partial harvesting also reduced the number of sprouts per stump for the red oak group and red maple. Sprouting probabilities were generally less in the Appalachian Plateau than the Ridge and Valley, particularly for the oaks (Quercus spp.). Partial harvesting systems decreased sprouting in both physiographic provinces. However, the sprouting in specific species groups varied between the two physiographic provinces. In the Ridge and Valley, the highest sprouting rates were in the clearcut for the red oak (60%), chestnut oak (77%), white oak-hickory (26%), and midstory (33%) species groups. Red maple sprouting was highest in the leave-tree (67%) in the Ridge and Valley. The mixed mesic and midstory groups were only reduced in the Ridge and Valley. Sprouting was negatively correlated with residual basal area for the red oak group, chestnut oak, and red maple. For the all oak species except white oak, sprouting was reduced by about 2% for every 1m²/ha increase in residual basal area.

A new arenicolous Boletus from the Gulf Coast of northern Florida.
About this Resource: Boletus abruptibulbus sp. nov. is described as new to science. It inhabits old sand dunes in mixed oak-pine stands across the Gulf Coast of northern Florida.

Modeling and measuring the effects of disturbance history an climate on carbon and water budgets in evergreen needleleaf forests
About this Resource: The effects of disturbance history, climate, and changes in atmospheric carbon dioxide (CO2) concentration and nitrogen deposition (N(dep)) on carbon and water fluxes in seven North American evergreen forests are assessed using a coupled water-carbon-nitrogen model, canopy-scale flux observations, and descriptions of the vegetation type, management practices, and disturbance histories at each site. The effects of interannual climate variability, disturbance history, and vegetation ecophysiology on carbon and water fluxes and storage are integrated by the ecosystem process model Biome-BGC, with results compared to site biometric analyses and eddy covariance observations aggregated by month and year. Model results suggest that variation between sites in net ecosystem carbon exchange (NEE) is largely a function of disturbance history, with important secondary effects from site climate, vegetation ecophysiology, and changing atmospheric CO2 and N(dep). The timing and magnitude of fluxes following disturbance depend on disturbance type and intensity, and on post-harvest management treatments such as burning, fertilization and replanting. The modeled effects of increasing atmospheric CO2 on NEE are generally limited by N availability, but are greatly increased following disturbance due to increased N mineralization and reduced plant N demand. Modeled rates of carbon sequestration over the past 200 years are driven by the rate of change in CO2 concentration for old sites experiencing low rates of N(dep). The model produced good estimates of between-site variation in leaf area index, with mixed performance for between- and within-site variation in evapotranspiration. There is a model bias toward smaller annual carbon sinks at five sites, with a seasonal model bias toward smaller warm-season sink strength at all sites. Various lines of reasoning are explored to help to explain these differences.

Interspecific divergence in foliar nutrient dynamics and stem growth in a temperate forest in response to chronic nitrogen inputs.
About this Resource: We studied the effects of excessive nitrogen (N) fertilization on foliar nutrient dynamics and stem growth in three important tree species in a mixed-deciduous forest. Stem diameter growth, foliar N concentrations, nitrogen-phosphorus (N/P) ratios, and nutrient resorption were determined for Acer rubrum L. (ACRU), Liriodendron tulipifera L. (LITU), and Prunus serotina Ehrh. (PRSE) on two 30-year-old watersheds at the Fernow Experimental Forest, West Virginia, USA: WS3, fertilized annually with 35 kg ammonium sulfate.ha(-1) since 1989, and WS7, an untreated control watershed. In an earlier (1992) study, foliar N concentrations of all three species averaged 11% higher in WS3 than in WS7. By 2000, that was no longer the case for any species; indeed N in ACRU leaves was 13% lower in WS3 that year. N/P ratios were elevated in WS3 only in PRSE in 1992 and in both ACRU and PRSE in 1997, but by 2001, mean N/P for all three species was lower in WS3. N resorption efficiencies were 30% lower in WS3 in ACRU and PRSE, but not in LITU. Stem diameter growth in WS3 was 55% lower in ACRU and 30% lower in LITU and PRSE compared with that in WS7. Results may indicate declining growth vigor in ACRU and, to a lesser extent, PRSE and LITU in the fertilized watershed. Observed interspecific differences in growth and plant nutrition responses suggest eventual changes in species composition under increasing N saturation.

Five-year radial growth of red oaks in mixed bottomland hardwood stands
About this Resource: We studied the relationships among 5-year radial (diameter and basal area) growth of red oak (genus Quercus, subgenus Erythrobalanus) crop trees and predictor variables representing individual tree vigor, distance-dependant competition measures, and distance-independent competition measures. The red oaks we examined are representative of the commercially and ecologically important oak species of the bottomland hardwood forests of the southeastern US. The crown class score, a quantitative measure of crown class and tree vigor, performed best in accounting for the variability in tree diameter growth. Plot-level variables failed to account for a significant proportion of the variability in tree radial growth. The basal area of the first-order neighbors that were taller than the crop trees and located within 2.4 times the mean overstory crown radius had the highest negative correlation with crop tree 5-year radial growth. Red oaks were a major part of these competitors and likely exerted the greatest competitive pressure. However, crop tree radial growth was positively associated with the basal area of the red oaks which were indirect (second order) neighbors and which were taller than the crop trees. It is possible that indirect neighbors do not compete with the crop trees, but they likely compete with the direct competitors of the crop trees, thus having an indirect positive influence on crop tree growth. Such reasoning is consistent with previously observed spatial dependence up to four times the mean overstory crown radius. The findings may have implications for thinning hardwoods stands and crop tree management in that foresters need to take into account (1) oak intra-genus competition, (2) the negative competitive effect of direct neighbors, and (3) the potentially positive effect of the indirect neighbors, the competitors' competitors.

Reproductive Responses of Northern Goshawks to Variable Prey Populations
About this Resource: Developing comprehensive conservation strategies requires knowledge of factors influencing population growth and persistence. Although variable prey resources are often associated with fluctuations in raptor demographic parameters, the mechanisms of food limitation are poorly understood, especially for a generalist predator like the northern goshawk (Accipiter gentilis). To determine the reproductive responses of goshawks to variable prey populations, we evaluated 823 goshawk breeding opportunities on the Kaibab Plateau, Arizona, USA, during 1994-2002. Concurrently, density was estimated for 4 prey species (2 avian, 2 mammalian). We explored the relationship between goshawk reproduction and prey density at one temporal scale (year) and 2 spatial scales (study area, forest type). Prey density for all 4 species combined accounted for 89% of the variation in goshawk reproduction within the entire study area (P < 0.001), 74% in mixed conifer forest (P = 0.003) and 85% in ponderosa pine (Pinus ponderosa) forest (P < 0.001). We found that an incremental increase in prey density resulted in a greater increase in goshawk reproduction in ponderosa pine forest than in mixed conifer forest, suggesting that the denser structural conditions of mixed conifer forest may have reduced prey availability. Red squirrel (Tamiasciurus hudsonicus) density explained more annual variation in goshawk reproduction within the study area (r2 = 0.87, P < 0.001), mixed conifer forest (r2 = 0.80, P = 0.001), and ponderosa pine forest (r2 = 0.85, P < 0.001) than did any other individual species. Although certain prey species were more strongly correlated with fluctuations in goshawk reproduction than were others, the high model selection uncertainty and the strong relationship between total prey density and number of goshawk fledglings produced indicated that alternate prey species were readily substituted for one another. Therefore, conservation strategies concerned with the status of goshawk populations should incorporate forest management practices that increase the abundance, diversity, and availability of prey resources.

Habitat Selection by American Martens in Coastal California
About this Resource: We investigated habitat selection using single- and mixed-scale modeling at 2 spatial scales, stand and home range, by the only known population of American martens (Martes americana) remaining in the historical range of the Humboldt subspecies (M. a. humboldtensis) in California, USA. During 2000 and 2001, we sampled a 12 x 14 grid with 2-km spacing, using 2 sooted track plates at each grid point. We detected martens at 26 of the 159 grid points. We used resource selection probability functions and an information-theoretic method to model habitat at detection locations. At the stand scale, martens selected conifer-dominated stands with dense, spatially extensive shrub cover (average = 74% cover, SE = 4) in the oldest developmental stage. At the home-range scale, martens selected the largest available patches ( average = 181 ha, SE = 14) of old-growth, old-growth and late-mature, or serpentine habitat. Mixed-scale models revealed that habitat characteristics from both scales best explained marten occurrence compared to one scale alone. Dense, spatially extensive shrub cover is a key habitat element for martens in coastal forests. Dense shrubs provide refuge from predators, cover for prey, and may also deter larger-bodied competitors. Managers can increase the likelihood of marten population persistence and encourage expansion in coastal forests by maintaining and restoring late-mature and old-growth, conifer-dominated forests with dense shrub cover in large, contiguous patches.

Initial tree regeneration responses to fire and thinning treatments in a Sierra Nevada mixed-conifer forest, USA
About this Resource: Fire is a driver of ecosystem patterns and processes in forests globally, but natural fire regimes have often been altered by decades of active fire management. Following almost a century of fire suppression, many western U.S. forests have greater fuel levels, higher tree densities, and are now dominated by fire-sensitive, shade-tolerant species. These fuel-loaded conditions can often result in high-intensity crown fires replacing historic low- to moderate-intensity fire regimes. In the mixed-conifer forests of the California Sierra Nevada, thinning and prescribed fire are widely used to reduce fuels and shift future stand composition from shade-tolerant species to more fire-resistant pines (Pinus lambertiana and Pinus jeffreyi) that were historically more abundant. The impacts of these treatments, however, on forest regeneration composition and abundance are unclear. We examined the effects of prescribed fire and common thinning treatments (understory and overstory thinning) on microsite conditions, seed rain, and tree regeneration in an old-growth, mixed-conifer forest in the Sierra Nevada, California, USA. Treatments significantly altered environmental conditions, but there was substantial variation and overlap in conditions among treatments. Seed rain of shade-tolerant Abies concolor and Calocedrus decurrens was 5-26 times greater than P. jeffreyi and P. lambertiana, creating inertia in efforts to shift stands towards increased pine abundance. Survival of Pinus germinants was greatest in burned microsites. The burn-overstory thin treatment had both the highest mortality of advanced A. concolor and C. decurrens regeneration and the greatest increase in pine regeneration. Species occupied microsites gradating from low light/high moisture to high light/low moisture in the order: C. decurrens, A. concolor, P. lambertiana, and P. jeffreyi. Results suggest prescriptions may need to thin mature A. concolor and C. decurrens to significantly reduce their seed rain, create an abundance of burned open microsites, or plant Pinus seedlings to shift regeneration composition in treated stands.

Factors Associated with Succession of Abandoned Agricultural Lands along the Lower Missouri River, U.S.A
About this Resource: The 1993 flood of the Missouri River led to the abandonment of agriculture on considerable land in the floodplain. This abandonment led to a restoration opportunity for the U.S. Federal Government, purchasing those lands being sold by farmers. Restoration of this floodplain is complicated, however, by an imperfect understanding of its past environmental and vegetative conditions. We examined environmental conditions associated with the current placement of young forests and wet prairies as a guide to the potential successional trajectory for abandoned agricultural land subject to flooding. We used Bayesian mixed-effects logistic regression to examine the effects of flood frequency, soil drainage, distance from the main channel, and elevation on whether a site was in wet prairie or in forest. Study site was included as a random effect, controlling for site-specific differences not measured in our study. We found, after controlling for the effect of site, that early-successional forest sites were closer to the river and at a lower elevation but occurred on drier soils than wet prairie. In a regulated river such as the lower Missouri River, wet prairie sites are relatively isolated from the main channel compared to early-successional forest, despite occurring on relatively moister soils. The modeled results from this study may be used to predict the potential successional fate of the acquired agricultural lands, and along with information on wildlife assemblages associated with wet prairie and forest can be used to predict potential benefit of these acquisitions to wildlife conservation.

Initial tree mortality and insect and pathogen response to fire and thinning restoration treatments in an old-growth mixed-conifer forest of the Sierra Nevada, California
About this Resource: Fire and thinning restoration treatments in fire-suppressed forests often damage or stress leave trees, altering pathogen and insect affects. We compared types of insect- and pathogen-mediated mortality on mixed-conifer trees 3 years after treatment. The number of bark beetle attacked trees was greater in burn treatments compared with no-burn treatments, and in some cases, larger pine trees were preferentially attacked. Restoration treatments are not expected to change the trajectory of spread and intensification of dwarf mistletoe. Thinning treatments may have provided a sanitation effect in which large leave trees have lower levels of dwarf mistletoe. Although thinning treatments are known to exacerbate root disease, <12% of cut stumps were infected with root pathogens (Armillaria gallica and Heterobasidion annosum). Treatments increased Ribes (alternate host for white pine blister rust) frequency and abundance, which may have very localized impacts on white pine blister rust dynamics. In some instances, fire, insects, and pathogens appear to conflict with forest restoration goals by reducing the percentage of pine and producing proportionally higher rates of tree mortality in large-diameter size classes. To better understand the long-term effects of restoration treatments on pathogens and insects, continued monitoring over the course of varying climatic conditions will be needed.

Forest composition following overstory mortality from southern pine beetle and associated treatments
About this Resource: Bark beetle caused mortality continues to play a critical role in determining the composition and structure of forests in North America, and revegetation dynamics following these disturbances are poorly understood. We assessed forest composition following southern pine beetle, Dendroctonus frontalis Zimmermann, mortality and associated cut and leave suppression, and compared them with undisturbed loblolly pine, Pinus taeda L., stands in Texas. Abundant hardwood regeneration dominated the understory, with little loblolly pine regeneration following either mortality event. Disturbances eliminated loblolly pine dominance in these even-aged stands, thus stratifying forest structure and apparently increasing stem density (stems·ha-1), richness (species·ha-1), and diversity (ha) in the lower strata. Aspect and elevation, presumably through influences on site moisture, were the primary gradients associated with vegetation variation in the canonical correspondence analyses for new regeneration. Mortality from Dendroctonus and cut and leave practices shifted loblolly pine communities to mixed upland hardwoods in model predictions generated by the southern variant of the Forest Vegetation Simulator. In addition to being an effective bark beetle control, cut and leave suppression did not alter predicted forest composition 50 years hence when compared with unsuppressed bark beetle-caused mortality. Because of the predicted shift to hardwood domination and a low pine basal area, it is expected that mortality from D. frontalis and cut and leave will substantially reduce future hazards from D. frontalis outbreaks.

Conifer regeneration in stand-replacement portions of a large mixed-severity wildfire in the Klamath-Siskiyou Mountains
About this Resource: Large-scale wildfires (approximately 10(4)-10(6) ha) have the potential to eliminate seed sources over broad areas and thus may lead to qualitatively different regeneration dynamics than in small burns; however, regeneration after such events has received little study in temperate forests. Following a 200 000 ha mixed-severity wildfire in Oregon, USA, we quantified (1) conifer and broadleaf regeneration in stand-replacement patches 2 and 4 years postfire; and (2) the relative importance of isolation from seed sources (live trees) versus local site conditions in controlling regeneration. Patch-scale conifer regeneration density (72%-80% Douglas-fir (Pseudotsuga menziesii (Mirb). Franco)) varied widely, from 127 to 6494 stems·ha-1. Median densities were 1721 and 1603 stems·ha-1 2 and 4 years postfire, respectively, i.e., approximately 12 times prefire overstory densities (134 stems·ha-1). Because of the complex burn mosaic, approximately 58% of stand-replacement area was <or=200 m from a live-tree edge (seed source), and approximately 81% was <or=400 m. Median conifer density exceeded 1000 stems·ha-1 out to a distance of 400 m from an edge before declining farther away. The strongest controls on regeneration were distance to live trees and soil parent material, with skeletal coarse-grained soils supporting lower densities (133 stems·ha-1) than fine-grained soils (729-1492 stems·ha-1). Other site factors (e.g., topography, broadleaf cover) had little association with conifer regeneration. The mixed-severity fire pattern strongly influenced the regeneration process by providing seed sources throughout much of the burned landscape.

Substrate pH Affects Nutrient Availability in Fertilized Douglas Fir Bark Substrates.
About this Resource: An experiment was conducted to determine how pH and nutrient availability in douglas fir bark (DFB) substrates respond to lime and sulfur (S) rates. The treatment design was a two-by-nine factorial arrangement with two substrate types and nine pH-altering amendments. The two substrates were 100% DFB or 75 DFB:15 sphagnum peatmoss:10 pumice (by volume). Substrate pH-altering amendments included elemental S amended at either 0.6 or 2.4 kg·m-3; calcium carbonate amended at 0.6, 1.5, and 5.9 kg·m-3; calcium hydroxide amended at 4.4, 8.9, or 23.7 kg·m-3; and a nonamended control. All substrates were amended by incorporating 0.9 kg·m-3 Micromax micronutrients before potting and topdressing 8 g/pot of 14N-4.2P-11.6K Osmocote controlled-release fertilizer after potting. A group of controls was also maintained for each substrate that received no fertilizer amendment (no S, lime, Micromax, or Osmocote). Four containers of each treatment were randomly selected and harvested 4 and 8 weeks after potting. Amendment with S decreased pH with increasing rate, whereas both lime types increased pH with increasing rate. The two substrates in general responded similarly to S and lime amendments, although there were some significant effects and interactions caused by substrate type. Ammonium-N and NO3-N both decreased exponentially with increasing substrate pH, whereas water-extractable phosphorus decreased linearly with increasing pH. Water-extractable potassium, calcium, magnesium, and sodium responded quadratically to increasing pH by initially decreasing and then increasing. The micronutrients boron and iron decreased with increasing pH, whereas DTPA extractions of manganese, zinc, and copper initially increased and then decreased over the range of observed pH.

Ectomycorrhizal characterization of an American chestnut ( Castanea dentata) -dominated community in Western Wisconsin
About this Resource: Circa 1900, a farmer from the eastern US planted 11 American chestnut (Castanea dentata) seeds on a newly established farm near West Salem in western Wisconsin. These trees were very successful, producing a large stand of over 6,000 trees. Since this area is well outside the natural range of chestnut, these trees remained free from chestnut blight until 1987. In the West Salem stand, chestnuts are the dominant species of a mixed forest community, reminiscent of the chestnut-oak ecosystems of pre-1900 Appalachia. To identify putative mycorrhizal associates of chestnut in this unique forest, our approach was twofold: (1) an extensive fruiting body survey was conducted for four seasons that yielded approximately 100 putative mycorrhizal species and (2) a belowground molecular approach was used to generate DNA sequences of the internal transcribed spacer region from ectomycorrhizae. Unexpectedly, chestnut did not appear to be the dominant underground ectomycorrhizal-forming plant species. This study highlights the need to identify the plant host species when conducting belowground molecular-based surveys and provides preliminary identification of ectomycorrhizal fungi associated with a disjunct stand of American chestnut.

Field efficiency and bias of snag inventory methods.
About this Resource: Snags and cavity trees are important components of forests, but can be difficult to inventory precisely and are not always included in inventories because of limited resources. We tested the application of N-tree distance sampling as a time-saving snag sampling method and compared N-tree distance sampling to fixed-area sampling and modified horizontal line sampling in mixed pine-hardwood forests of southern Maine and New Hampshire. We also present a novel modification of N-tree distance sampling that limits the distance from plot center that an observer must search to find tally trees. A field test shows N-tree to be quick, but generally biased and characterized by high variability. Distance-limited N-tree sampling mitigates these problems, but not completely. We give recommendations for operational snag inventory in similar forest types.

Determinants of Timberland Use by Ownership and Forest Type in Alabama and Georgia
About this Resource: Land use changes and timberland use by ownership and forest type in Alabama and Georgia between 1972 and 2000 are analyzed using a modified multinomial logit approach. Low average land quality, federal cost-share incentives, and favorable returns to forestry relative to agriculture were the main factors associated with timberland increase. Higher forestry returns helped increase industrial timberland but not nonindustrial private forests. An increase in hardwood forests at the expense of softwood and mixed forests was driven by increasing hardwood returns. Increasing softwood returns and tree planting assistance programs alleviated declines in softwood forests. Because factors influencing timberland use changes differ by ownership and forest type, treating all timberland as one major category may lead to incorrect predications.

Fruit removal of a wild tomato, Solanum granulosoleprosum Dunal (Solanaceae), by birds, bats and non-flying mammals in an urban Brazilian environment
About this Resource: A study of removal of fruits of the wild tomato, Solanum granulosoleprosum Dunal (N = 5 plants), by vertebrates was carried out in an urban environment of southern Brazil from January to May 1997 and February 1998. To verify diurnal and nocturnal removals, fruits were counted in several fruit bunches, being classified by size and color. Diurnal observations were made on plants to verify bird removal. A mist net was placed among the plants from the evening to 23:00 h to verify bat consumption. Live traps baited with S. granulosoleprosum fruits were placed on the ground among plants to verify terrestrial removers. On average it was found two ripe fruits available per bunch/day, but unripe, small, fruits were dominant (70%). Nocturnal mammals and birds-diurnal mammals partitioned fruits similarly. Bats removing fruits were Artibeus lituratus (Olfers, 1818), Pygoderma bilabiatum (Wagner, 1843) and Sturnira lilium (E. Geoffroy, 1810). Birds were Saltator similis Lafresnaye & d'Orbigny, 1837 and Thraupis sayaca (Linnaeus, 1766). Terrestrial mammals were a marsupial and three rodent species. Except for rodents, these vertebrates must be promoting the seed dispersal of S. granulosoleprosum seeds in disturbed mixed forests of southern Brazil.

VEMAP vs VINCERA: A DGVM sensitivity to differences in climate scenarios
About this Resource: The MC1 DGVM has been used in two international model comparison projects, VEMAP (Vegetation Ecosystem Modeling and Analysis Project) and VINCERA (Vulnerability and Impacts of North American forests to Climate Change: Ecosystem Responses and Adaptation). The latest version of MC1 was run on both VINCERA and VEMAP climate and soil input data to document how a change in the inputs can affect model outcome. We compared simulation results under the two sets of future climate scenarios and reported on how the different inputs can affect vegetation distribution and carbon budget projections. Under all future scenarios, the interior West becomes woodier as warmer temperatures and available moisture allow trees to get established in grasslands areas. Concurrently, warmer and drier weather causes the eastern deciduous and mixed forests to shift to a more open canopy woodland or savanna type while boreal forests disappear almost entirely from the Great Lakes area by the end of the 21st century. While under VEMAP scenarios the model simulated large increases in carbon storage in a future woodier West, the drier VINCERA scenarios accounted for large carbon losses in the east and only moderate gains in the West. But under all future climate scenarios, the total area burned by wildfires increased especially in C4 grasslands under all scenarios and in dry woodlands under VINCERA scenarios. The model simulated non-agricultural lands in the conterminous United States as a source of carbon in the 21st century under the VINCERA future climate scenarios but not VEMAP. However, the magnitude of this carbon source to the atmosphere could be greatly reduced if the CO2 growth enhancement factor built in the model was enhanced but evidence that all mature forests across the entire country will respond positively to increased atmospheric CO2 is still lacking.

Effect of Ecological Aspects on the Quantitative Variables of Mixed Broadleaf Forests Using Multivariate Statistical Method
About this Resource: This research was performed on the commercial forests of the western Caspian region, Shafa-Rud basin, to investigate firstly the effect of ecological aspects (northern and southern) on the quantitative variables of mixed broadleaf forests using a multivariate statistical method (MANOVA). The second aim was to estimate the volume growth of Shafa-Rud forest basin. Two northern and southern aspects were chosen in the middle latitude profile (800-1200 m.a.s.l) and on each aspect, 30 circular sampling plots were selected using the random blocking method. In each sample plot, six trees were chosen according to a predefined model for sampling the radial growth using an increment borer and for measuring other quantitative variables such as height of trees and diameter at breast height (DBH). Weighted local volume tables were calculated for each of the ecological aspects and were compared using t-test. Multivariate statistical analysis, Hotelling`s T22 test, which was carried out on means vector of the four quantitative variables: diameter, height, bark thickness and radial growth, demonstrated that the difference in centroid of vectors was highly significant. The stand volume growths in the southern and northern aspects were found to be 9.2 and 9.5 sylve per hectare, respectively.

Stand Structure and Spatial Patterns of Trees in Mixed Hyrcanian Beech Forest, Iran
About this Resource: The mixed beach forests (Fagus orientalis) commonly dominate by shade tolerance species with irregular uneven age stand structure. The aim of this study was to analyze the stand structure and spatial pattern in order to identify specific structural patterns. Data was collected from a 16 ha permanent plot. We mapped all stems >7.5 cm in diameter at breast height on permanent plot. The six main species were divided into two groups based on density and stand structure. Group A had higher density than group B, as well as L-shaped DBH distribution of live stems. Species in group B had bell-shaped DBH distributions. Species in group A have clump spatial distribution pattern in all layers but clump intensity is more than in understory layer and size of patch clump is small in this group. This phenomenon for group A may explaining by having numerous coppice, sucker and patch regeneration in the understory layer. Middlestory and understory stems of the six major tree species were patchily distributed throughout the plot but for Alder and Maple species common pattern in canopy layer was complete spatial randomness. The distribution of Beech and Hornbeam trees were negatively associated with other species. These results suggest species differences in favorable canopy condition.

Band-tailed pigeon distribution and habitat component availability in Western Oregon
About this Resource: The abundance of Pacific coast band-tailed pigeons (Columba fasciata monilis) has declined; this decline is thought to be due, in part, to habitat alteration associated with current forestry practices which replace diverse forest stands with even-aged conifer monocultures and control understorey shrubs and food availability. We examined distribution of Pacific coast band-tailed pigeons and availability of habitat components (known mineral sites and potential nesting and feeding areas) in the central Coast Range of Oregon, USA. Also, we evaluated associations of relative pigeon abundance at 323 randomly selected sample points and 97 known nest sites with adjacent habitat component availability. In Oregon, the pigeons nest primarily in Douglas fir (Pseudotsuga menziesii) trees in closed-canopy conifer or mixed forests and feed on berries of red elder (Sambucus racemosa), blue elder (S. cerulea) and cascara (Rhamnus purshiana). We detected 1-8 pigeons during a 1-hour point count at 83% of sample points from mid-June to July 1996-98. Eighty-four percent of the study area was classified as potential nesting (66%) and feeding (18%) areas. The maximum distance to the closest known mineral site was 37 km, but 91% of the study area was within 25 km of the nearest known mineral site. Availability of habitat components around known nest sites was similar to that at sample points and reflected availability within the study area. Relative pigeon abundance was not associated with availability of known mineral sites, or potential nesting and feeding areas. Pigeons were 62% more abundant along the western one-third of the Coast Range than along the eastern one-third, however, and this pattern was correlated with the apparent availability of red elder and cascara . Possibly the availability of red elder and cascara imposed a constraint on band-tailed pigeon distribution and abundance.

Characterisation of ecosystem water-use efficiency of european forests from eddy covariance measurements
About this Resource: Water-use efficiency (WUE) has been recognized as an important characteristic of vegetation productivity in various natural scientific disciplines for decades, but only recently at the ecosystem level, where different ways exist to characterize water-use efficiency. Hence, the objective of this research was (a) to systematically compare different ways of calculating ecosystem water-use efficiency (WUEe) from eddy-covariance measurements, (b) quantify the diurnal, seasonal and interannual variability of WUEe in relation to meteorological conditions, and (c) analyse between-site variability of WUEe as affected by vegetation type and climatic conditions, across sites in European forest ecosystems. &lt;br&gt;&lt;br&gt; Day-to-day variability of gross primary productivity (GPP) and evapotranspiration (ET) were more strongly coupled than net ecosystem production (NEP) and ET, obviously because NEP also depends on the respiration that is not heavily coupled to water fluxes. However, the slope of daytime NEP versus ET (m&lt;sub&gt;NEP&lt;/sub&gt;) from half-hourly measurements of a single day may also be used as a WUEe-estimate giving very similar results to those of the GPP-ET slope (m&lt;sub&gt;GPP&lt;/sub&gt;), since the diurnal variation is dominated by GPP. Since ET is the sum of transpiration (linked to GPP) and evaporation from wet vegetation and soil surfaces (not linked to GPP) we expected that WUEe is increasing when days after rain are excluded from the analysis. However only very minor changes were found, justifying an analysis of WUEe related to vegetation type. &lt;br&gt;&lt;br&gt; In most of the studied ecosystems the instantaneous WUE&lt;sub&gt;GPP&lt;/sub&gt; was quite sensitive to diurnally varying meteorological conditions and tended to decline from the morning to the afternoon by more than 50% because of increasing vapour pressure deficits (VPD). &lt;br&gt;&lt;br&gt; Seasonally, WUE&lt;sub&gt;GPP&lt;/sub&gt; increased with a rising monthly precipitation sum and rising average monthly temperatures up to a threshold of 11, 14 and 18&deg;C in boreal, temperate and Mediterranean ecosystems, respectively. Across all sites, the highest monthly WUE&lt;sub&gt;GPP&lt;/sub&gt;-values were detected at times of positive anomalies of summer-precipitation. During drought periods with high temperatures, high VPD, little precipitation and low soil water content, the water-use efficiency of gross carbon uptake (WUE&lt;sub&gt;GPP&lt;/sub&gt;) tended to decrease in all forest types because of a stronger decline of GPP compared to ET. &lt;br&gt;&lt;br&gt; However the largest variation of growing season WUE&lt;sub&gt;GPP&lt;/sub&gt; was found between-sites and significantly related to vegetation type: WUE&lt;sub&gt;GPP&lt;/sub&gt; was highest in ecosystems dominated by deciduous trees ranging from 5.0 g CO&lt;sub&gt;2&lt;/sub&gt; kg H&lt;sub&gt;2&lt;/sub&gt;O&lt;sup&gt;&minus;1&lt;/sup&gt; for temperate broad-leaved deciduous forests (TD), to 4.5 for temperate mixed forests (TM), 3.5 for temperate evergreen conifers (TC), 3.4 for Mediterranean broad-leaved deciduous forests (MD), 3.3 for Mediterranean broad-leaved evergreen forests (Mbeg), 3.1 for Mediterranean evergreen conifers (MC), 2.9 for boreal evergreen conifers (BC) and only 1.2 g CO&lt;sub&gt;2&lt;/sub&gt; kg H&lt;sub&gt;2&lt;/sub&gt;O&lt;sup&gt;&minus;1&lt;/sup&gt; for a boreal wetland site (BT). Although vegetation type and meteorology co-vary, the WUEGPP variation was hardly related to meteorology, as we could show by comparing similar meteorological conditions only. Furthermore we compared across-site WUE&lt;sub&gt;GPP&lt;/sub&gt; only under conditions when the 10% high GPP rates were exhibited. The between site differences remained, and at all sites ecosystem reached higher WUE&lt;sub&gt;GPP&lt;/sub&gt; levels under this condition. This means when vegetation is most productive usually it also maximises the amount of carbon gained per water lost. &lt;br&gt;&lt;br&gt; Overall our results show that water-use efficiency exhibits a strong time-scale dependency in the sense that at longer time-scale meteorological conditions play a smaller role compared to shorter time scale. Moreover, we highlight the role of vegetation in determining carbon-water relation at ecosystem level. Consequently, all predictions of changing carbon-water cycle under changing climate should take into this role and the differences between vegetation types. These results show the strong time-scale dependency of water-use efficiency

Influences of climate on fire regimes in montane forests of north-western Mexico
About this Resource: To identify the influence of interannual and interdecadal climate variation on the occurrence and extent of fires in montane conifer forests of north-western Mexico. This study was conducted in Jeffrey pine (Pinus jeffreyi Grev. & Balf.)-dominated mixed-conifer forests in the central and northern plateau of the Sierra San Pedro Mártir, Baja California, Mexico. Fire occurrence was reconstructed for 12 dispersed sites for a 290-year period (1700-1990) from cross-dated fire-scarred samples extracted from live trees, snags and logs. Superposed epoch analysis was used to examine the relationships of tree-ring reconstructions of drought, the El Niño/Southern Oscillation (ENSO) and the Pacific Decadal Oscillation (PDO) with fire occurrence and extent. Years with no recorded fire scars were wetter than average. In contrast, years of widespread fires were dry and associated with phase changes of the PDO, usually from positive (warm) to negative (cold). The influence of the PDO was most evident during the La Niña phase of the ENSO. Widespread fires were also associated with warm/wet conditions 5 years before the fire. We hypothesize that the 5-year lag between warm/wet conditions and widespread fires may be associated with the time necessary to build up sufficient quantity and continuity of needle litter to support widespread fires. Two periods of unusually high fire activity (1770-1800 and 1920-1950) were each followed by several decades of unusually low fire activity. The switch in each case was associated with strong phase changes in both PDO and ENSO. Climate strongly influences fire regimes in the mountains of north-western Mexico. Wet/warm years are associated with little fire activity. However, these years may contribute to subsequent fire years by encouraging the production of sufficient needle litter to support more widespread fires that occur in dry/cool years.

Spatially explicit modeling of mixed-severity fire regimes and landscape dynamics
About this Resource: Simulation models of disturbance and succession are being increasingly applied to characterize landscape composition and dynamics under natural fire regimes, and to evaluate alternative management strategies for ecological restoration and fire hazard reduction. However, we have a limited understanding of how landscapes respond to changes in fire frequency, and about the sensitivity of model predictions to assumptions about successional pathways and fire behavior. We updated an existing landscape dynamics model (LADS) to simulate the complex interactions between forest dynamics, fire spread, and fire effects in dry forests of the interior Pacific Northwest. Experimental model runs were conducted on a hypothetical landscape at fire rotations ranging from 5 to 50 years. Three sensitivity analyses were carried out to explore the responses of landscape composition to (1) parameters characterizing succession and fire effects on vegetation, (2) the probability of fire spread into different successional stages, and (3) the size and spatial pattern of static fire refugia. The area of old open-canopy forests was highest at the shortest fire rotations, and was particularly sensitive to the probability of stand-replacement fire in open-canopy forests and to the fire-free period required for ingrowth to occur in open-canopy forests. The area of old closed-canopy forests increased with lengthening fire rotation, but always comprised a relatively small portion of the landscape (<10%). The area of old closed-canopy forests increased when fire spread was more rapid in open-canopy forests than in closed-canopy forests, and when the physical landscape incorporated large “fire refugia” with low fire spread rates. Old closed-canopy forests appear to comprise a relatively minor landscape component in mixed-severity fire regimes with fire rotations of 50 years or less. However, these results are sensitive to assumptions about the spatial interactions between fire spread, landscape vegetation patterns, and the underlying physical landscape.

Distribution and abundance of Neotoma in western Oregon and Washington
About this Resource: Bushy-tailed woodrats (Neotoma cinerea) and dusky-footed woodrats (N. fuscipes) add substantially to the prey base of many avian and mammalian predators. High biomass of woodrats can reduce markedly area requirements of predators; thus, management for woodrats has potential in conservation. But patterns of abundance of woodrats in the Pacific Northwest are poorly understood. Our objective was to determine local abundances and regional distributions of N. cinerea and N. fuscipes in forests west of the Crest of the Cascade Range in Oregon and Washington. We sampled a variety of forests from 1985 to 1997 in seven physiographic provinces. In Washington, we found that N. cinerea was rare in upland forests, but abundant along rocky streams on the eastern Olympic Peninsula and in rock bluffs on the west slope of the Cascade Range; N. fuscipes does not occur in Washington. In Oregon, N. fuscipes is at the northern limits of its range and we found that it was rare in Douglas-fir (Pseudotsuga menziesii)-western hemlock (Tsuga heterophylla) forests; N. cinerea was common in old forests and along streams. In mixed-conifer forests both species occasionally were abundant, but their abundances were negatively correlated. Neotoma cinerea was rare in mixed-conifer-mixed-evergreen forests but N. fuscipes occasionally was abundant in early, and present in late stages of forest development. The distribution of N. fuscipes can be explained by its preference for dense shrub cover and it ability to consume plants potentially toxic to other mammals; the only compelling explanation for the irregular distribution of N. cinerea is exceptional vulnerability to predation because of its size and social behavior. Because of zoogeographic restrictions, limited opportunities exist in western Oregon and Washington to manage habitat for woodrats as a means of assisting in the recovery and maintenance of viable populations of predators sensitive to loss or management of forests